Har Gobind Khorana

Har Gobind Khorana
Born	(1922-01-09)9 January 1922 Raipur, Multan, Punjab Province, British India
Died	9 November 2011(2011-11-09) (aged 89) Concord, Massachusetts, U.S.
Citizenship	United States
Alma mater	Government College University, Lahore Punjab University (BS, MS) University of Liverpool (PhD)
Known for	First to demonstrate the role of nucleotides in protein synthesis
Spouse	Esther Elizabeth Sibler
Awards	Nobel Prize in Medicine (1968) Gairdner Foundation International Award (1980) Louisa Gross Horwitz Prize ForMemRS (1978) Albert Lasker Award for Basic Medical Research Padma Vibhushan Willard Gibbs Award (1974)
Scientific career
Fields	Molecular biology
Institutions	MIT (1970–2007) University of Wisconsin–Madison (1960–70) University of British Columbia (1952–60) University of Cambridge (1950–52) Swiss Federal Institute of Technology, Zurich (1948–49)
Doctoral advisor	Roger J.S. Beer
Doctoral students	Shiladitya DasSarma
Signature

Har Gobind Khorana (9 January 1922 – 9 November 2011) was an Indian-American biochemist.^[1] While on the faculty of the University of Wisconsin–Madison, he shared the 1968 Nobel Prize for Physiology or Medicine with Marshall W. Nirenberg and Robert W. Holley for research that showed the order of nucleotides in nucleic acids, which carry the genetic code of the cell and control the cell's synthesis of proteins. Khorana and Nirenberg were also awarded the Louisa Gross Horwitz Prize from Columbia University in the same year.^[2][3]

Born in British India, Khorana served on the faculties of three universities in North America. He became a naturalized citizen of the United States in 1966,^[4] and received the National Medal of Science in 1987.^[5]