Bryan Clarke

Bryan Clarke FRS
Born	Bryan Campbell Clarke (1932-06-24)24 June 1932 Gatley, Cheshire[2] (now Greater Manchester)
Died	27 February 2014(2014-02-27) (aged 81)
Alma mater	University of Oxford (BA, DPhil)
Spouse	Ann Clarke (née Jewkes)[2]
Children	One daughter, Alex; one son, Peter
Awards	Linnean Medal (2003) Darwin-Wallace Medal (2008) Darwin Medal (2010)
Scientific career
Fields	Genetics, evolutionary biology
Institutions	University of Nottingham University of Edinburgh
Thesis	Some factors affecting shell colour polymorphism in Cepaea (1961)
Doctoral advisor	Arthur Cain
Doctoral students	John Endler Steve Jones[1]
Other notable students	Fred W. Allendorf

Bryan Campbell Clarke FRS^[3] (24 June 1932 – 27 February 2014) was a British Professor of genetics, latterly emeritus at the University of Nottingham. Clarke is particularly noted for his work on apostatic selection (which is a term he coined in 1962) and other forms of frequency-dependent selection, and work on polymorphism in snails, much of it done during the 1960s. Later, he studied molecular evolution. He made the case for natural selection as an important factor in the maintenance of molecular variation, and in driving evolutionary changes in molecules through time. In doing so, he questioned the over-riding importance of random genetic drift advocated by King, Jukes, and Kimura. With Professor James J Murray Jnr (University of Virginia), he carried out an extensive series of studies on speciation in land snails of the genus Partula inhabiting the volcanic islands of the Eastern Pacific. These studies helped illuminate the genetic changes that take place during the origin of species.