Gamma diversity

In ecology, gamma diversity (γ-diversity) is the total species diversity in a landscape. The term was introduced by R. H. Whittaker^[1] together with the terms alpha diversity (α-diversity) and beta diversity (β-diversity). Whittaker's idea was that the total species diversity in a landscape (γ) is determined by two different things, the mean species diversity in sites at a more local scale (α) and the differentiation among those sites (β). According to this reasoning, alpha diversity and beta diversity constitute independent components of gamma diversity:

γ = α × β