Mandibulata

Mandibulata; Temporal range:
	The mandibles of a bull ant
	Life restoration of Odaraia, an early mandibulate belonging to Hymenocarina with mandibles present near the mouth
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Mandibulata; de Clairville, 1798
Subdivisions
incertae sedis
	†Acheronauta?; †Aquilonifer; †Bradoriida?; †Cambropachycopidae?; †Cambropodus?; †Captopodus?; †Cascolus?; †Fuxianhuiida; †Martinssonia?; †Rhyniognatha?; †Skaracarida?; †Tanazios?; †Thylacocephala?; †Oelandocarididae?; †Musacaris?;

The clade Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans, myriapods (centipedes and millipedes, among others), and all true insects. The name "Mandibulata" refers to the mandibles, a modified pair of limbs used in food processing, the presence of which are characteristic of most members of the group.

The mandibulates are divided between the extant groups Myriapoda (millipedes and centipedes, among others) and Pancrustacea (including crustaceans and hexapods, the latter group containing insects). Molecular phylogenetic studies suggest that the living arthropods are related as shown in the cladogram below. Crustaceans do not form a monophyletic group as insects and other hexapods have evolved from within them.^[1]^[2]^[3]

A number of extinct groups have also been placed in Mandibulata, including Hymenocarina,^[4] Euthycarcinoidea,^[5] and possibly Fuxianhuiida.^[6]

Arthropoda

Mandibulata

	Pancrustacea (crustaceans and hexapods)

	Myriapoda (centipedes, millipedes, and allies)

Chelicerata (sea spiders, horseshoe crabs, and arachnids)

^ Cite error: The named reference Regier was invoked but never defined (see the help page).
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^ Izquierdo-López, Alejandro; Caron, Jean-Bernard (August 2024). "The Cambrian Odaraia alata and the colonization of nektonic suspension-feeding niches by early mandibulates". Proceedings of the Royal Society B: Biological Sciences. 291 (2027). doi:10.1098/rspb.2024.0622. ISSN 1471-2954. PMC 11463219. PMID 39043240.
^ Edgecombe, Gregory D.; Strullu-Derrien, Christine; Góral, Tomasz; Hetherington, Alexander J.; Thompson, Christine; Koch, Marcus (2020). "Aquatic stem group myriapods close a gap between molecular divergence dates and terrestrial fossil record". Proceedings of the National Academy of Sciences. 117 (16): 8966–8972. doi:10.1073/pnas.1920733117. PMC 7183169. PMID 32253305. S2CID 215408474.
^ Aria, Cédric; Zhao, Fangchen; Zhu, Maoyan (September 2021). "Fuxianhuiids are mandibulates and share affinities with total-group Myriapoda". Journal of the Geological Society. 178 (5). doi:10.1144/jgs2020-246. ISSN 0016-7649.

[Regier-1] Cite error: The named reference Regier was invoked but never defined (see the help page).

[vonJennWillDell11-2] Cite error: The named reference vonJennWillDell11 was invoked but never defined (see the help page).

[RotaStabelli11-3] Cite error: The named reference RotaStabelli11 was invoked but never defined (see the help page).

[:8-4] Izquierdo-López, Alejandro; Caron, Jean-Bernard (August 2024). "The Cambrian Odaraia alata and the colonization of nektonic suspension-feeding niches by early mandibulates". Proceedings of the Royal Society B: Biological Sciences. 291 (2027). doi:10.1098/rspb.2024.0622. ISSN 1471-2954. PMC 11463219. PMID 39043240.

[5] Edgecombe, Gregory D.; Strullu-Derrien, Christine; Góral, Tomasz; Hetherington, Alexander J.; Thompson, Christine; Koch, Marcus (2020). "Aquatic stem group myriapods close a gap between molecular divergence dates and terrestrial fossil record". Proceedings of the National Academy of Sciences. 117 (16): 8966–8972. doi:10.1073/pnas.1920733117. PMC 7183169. PMID 32253305. S2CID 215408474.

[6] Aria, Cédric; Zhao, Fangchen; Zhu, Maoyan (September 2021). "Fuxianhuiids are mandibulates and share affinities with total-group Myriapoda". Journal of the Geological Society. 178 (5). doi:10.1144/jgs2020-246. ISSN 0016-7649.

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